CIESM Atlas of Exotic Molluscs in the Mediterranean

 
The data base of Exotic Molluscs in the Mediterranean Sea was developed by a CIESM task force of four biologists : Drs Argyro Zenetos (Natl. Center for Marine Research, Athens), Serge Gofas (Univ. Malaga), Giovanni Russo (Univ. di Napoli), Jose Templado (Museo Natl. de Ciencas Naturales, Madrid).
Updates are regularly provided by the team of experts now coordinated by Dr Gofas.
 



ERRATUM: In January 2005 a major update corrected a number of errors present in the original printed version. We are deeply grateful to Prof. Bella Galil (Israel Oceanographic and Limnological Research, Haifa), Drs Henk Mienis, Bogi Cesare, Giovanni Buzzurro, Anna Occhipinti-Ambrogi and Paul Clark for their contribution to the Atlas of Exotic Molluscs.
---------------------------------------------------------------------------------------


Check-list of exotic species 1 (total: 137)
1 Only those exotic species of Indo-Pacific origin that were recorded after 1920 and of Atlantic origin that were recorded after 1960 are considered.

Origin:
AA = American Atlantic, AS = Arabian Sea, CIRT = Circumtropical, IO = Indian Ocean, IP = Indo-Pacific, NEP = North Eastern Pacific Ocean, PG = Persian Gulf, PO = Pacific Ocean, RS = Red Sea, SC = Suez Canal, TP = Tropical Pacific Ocean, TA = Tropical Atlantic, WP = Western Pacific Ocean.

Status:
A = Alien: considered non-established, on the basis of one or two published records, often a single specimen.
E = Established: presumed established on the basis of at least two distincts records.
Q = Questionable: applies for a few taxa to which there remain unsettled taxonomic issues.

 
Taxon
Origin
Status
Last update
of the species sheet:
GASTROPODA - PROSOBRANCHIA
Nacellidae

Cellana rota
RS, IO
E
Dec 2003
Haliotidae (abalones)
Haliotis pustulata cruenta

RS

A
Dec 2003
Fissurellidae (keyhole limpets)
Diodora ruppellii

IP, RS

E
Dec 2003
Neritidae
Smaragdia souverbiana

Nerita sanguinolenta

IP, RS

RS

E

A

Dec 2003

Jan 2005
Trochidae (top shells)
Trochus erithraeus

Pseudominolia nedyma

Stomatella impertusa

RS

RS, IO

IP, RS

E

E

A

Jan 2005

Dec 2003

Dec 2003
Cerithiidae
Cerithium nesioticum

Cerithium scabridum

Cerithium egenum

Rhinoclavis kochi

Clypeomorus bifasciatus

RS, IP

RS, IO

IP, RS

IP, RS

RS, IP

A

E

A

E

A

Jan 2005

Jan 2005

Dec 2003

Jan 2005

Dec 2003
Planaxidae
Angiola punctostriata

Planaxis griseus

SC, IP

RS

A

A

Jan 2005

Jan 2005
Litiopidae
Gibborissoa virgata

IP

E
Dec 2003
Obtortionidae
Finella pupoides

Clathrofenella ferruginea

IP

IP, RS

E

E

Jan 2005

Jan 2005
Dialidae
Diala varia

IP, RS

E
Jan 2005
Cerithiopsidae
Cerithiopsis pulvis

Cerithiopsis tenthrenois

RS

IO

E

E

Jan 2005

Dec 2003
Triphoridae
Metaxia bacillum

RS

E
Jan 2005
Rissoidae
Rissoina bertholleti

Rissoina spirata

Alvania dorbignyi

Voorwindia tiberiana

RS, IO

IP

RS

RS

E

A

E

A

Jan 2005

Dec 2003

Dec 2003

Dec 2003
Strombidae (conchs)
Strombus persicus

Strombus mutabilis

PG, AS

IP, RS

E

A

Jan 2005

Dec 2003
Hipponicidae
Sabia conica

RS, IP

A
Dec 2003
Calyptraeidae (slipper shells)
Crepidula aculeata

Crepidula fornicata

AA

AA

E

E

Dec 2003

Dec 2003
Cypraeidae (cowries)
Erosaria turdus

Purpuradusta gracilis notata

Palmadusta lentiginosa

RS, IO

IO, RS

IO, PG

A

E

A

Dec 2003

Jan 2005

Dec 2003
Naticidae (moon shells)
Natica gualteriana

RS, IP

E
Dec 2003
Epitoniidae (wentle traps)
Cycloscala hyalina

RS, IP

E
Jan 2005
Eulimidae
Sticteulima cf. lentiginosa

IP

A
Jan 2005
Muricidae (murex shells)
Ergalatax obscura

Thais lacera

Thais sacellum

Rapana venosa

Murex forskoehlii

RS, AS

IO, PG

IO, RS

PO

RS, PG

E

E

E

E

E

Jan 2005

Jan 2005

Jan 2005

Dec 2003

Jan 2005
Nassariidae
Nassarius arcularia plicatus

RS, IO

A
Dec 2003
Fasciolariidae (spindle shells)
Fusinus verrucosus

RS

E
Jan 2005
Columbellidae (dove shells)
Zafra savignyi

Zafra selasphora

RS

IO, RS

E

E

Dec 2003

Jan 2005
Conidae (cone shells)
Conus fumigatus

RS

A
Dec 2003
GASTROPODA - HETEROBRANCHIA
Anisocyclidae
Murchisonella columna

IP

A
Dec 2003
Pyramidellidae
Chrysallida fischeri

Chrysallida maiae

Chrysallida pirintella

Adelactaeon fulvus

Adelactaeon amoenus

Styloptygma beatrix

Cingulina isseli

Turbonilla edgarii

Syrnola fasciata

Syrnola cinctella

Odostomia lorioli

Oscilla jocosa

Iolaea neofelixoides

Hinemoa cylindrica

Leucotina cfr. eva

RS

RS

RS

IP

IP

PG

RS

IP

IP

IP, RS

RS

RS, PG

PO

IP

IP

E

E

E

E

E

E

E

E

E

A

A

A

A

A

A

Dec 2003

Dec 2003

Jan 2005

Jan 2005

Dec 2003

Dec 2003

Dec 2003

Jan 2005

Dec 2003

Dec 2003

Dec 2003

Dec 2003

Dec 2003

Dec 2003

Dec 2003
GASTROPODA - OPISTHOBRANCHIA
Cylichnidae

Acteocina mucronata

RS

E
Dec 2003
Retusidae
Cylichnina girardi

Pyrunculus fourierii

IP

IP, RS

E

E

Jan 2005

Dec 2003
Bullidae (bubble shells)
Bulla ampulla

IP

E
Dec 2003
Haminoeidae
Haminoea callidegenita

Haminoea cyanomarginata

PO, AA

RS

E

A

Dec 2003

Dec 2003
Aglajidae
Chelidonura fulvipunctata

IP

E
Jan 2005
Aplysiidae (sea hares)
Bursatella leachi

CIRT

E
Dec 2003
Pleurobranchidae
Pleurobranchus forskalii

RS

A
Dec 2003
Polyceridae
Polycerella emertoni

Polycera hedgpethi

TA

NEP

E

A

Dec 2003

Dec 2003
Triophidae
Plocamopherus ocellatus

RS

E
Jan 2005
Discodorididae
Discodoris lilacina

IP, RS

E
Dec 2003
Chromodorididae
Hypselodoris infucata

Chromodoris quadricolor

IP, RS

RS

E

A

Dec 2003

Dec 2003
Dendrodorididae
Dendrodoris fumata

RS, IP

A
Dec 2003
Tethyidae
Melibe fimbriata

IO, PO

E
Jan 2005
Tergipedidae
Cuthona perca

PO

A
Dec 2003
Flabellinidae
Flabellina rubrolineata

IP, RS

E
Jan 2005
Facelinidae
Caloria indica

IP

A
Jan 2005
Aeolidiidae
Aeolidiella indica

CIRT

A
Dec 2003
GASTROPODA - PULMONATA
Siphonaridae
Siphonaria crenata

IP, RS

E
Dec 2003
BIVALVIA - PTEROMORPHIA
Arcidae (arc clams)
Acar plicata

Anadara demiri

Anadara inaequivalvis

Anadara natalensis

IP, RS

IO

IP

WIO, RS

A

E

E

E

Dec 2003

Dec 2003

Dec 2003

Jan 2005
Glycymeridae (bittersweet clams)
Glycymeris arabicus
WIO, RS
A
Dec 2003
Limopsidae
Limopsis multistriata

IP, RS

A
Dec 2003
Mytilidae (mussels)
Musculista perfragilis

Musculista senhousia

Modiolus auriculatus

Xenostrobus securis

Brachidontes pharaonis

Septifer forskali
IP, RS
WP

WI, RS

TP

IO, RS

RS
E
E

A

E

E

A

Jan 2005

Jan 2005

Jan 2005

Dec 2003

Jan 2005

Jan 2005
Ostreidae (oysters)
Crassostrea gigas

Saccostrea cucullata

Saccostrea commercialis

Dendrostrea frons

NWP

IO, RS

PO

IP

E

E

A

E

Dec 2003

Dec 2003

Dec 2003

Dec 2003
Pteriidae (pearl oysters)
Pinctada margaritifera

Pinctada radiata

IP, RS

IP, RS

E

E

Dec 2003

Jan 2005
Malleidae (hammer oysters)
Malvufundus regulus

IP, RS

E
Jan 2005
Pectinidae (scallops)
Chlamys lischkei

AA

A
Dec 2003
Spondylidae (spiny oysters)
Spondylus spinosus

Spondylus groschi

Spondylus cf. multisetosus

IP, RS

WIO, RS

IP

E

Q

Q

Jan 2005

Dec 2003

Dec 2003
BIVALVIA - HETERODONTA
Lucinidae (lucina clams)
Divalinga arabica

RS, AS

E
Jan 2005
Trapeziidae
Trapezium oblongum

IP, RS

A
Jan 2005
Ungulinidae
Diplodonta cf. subrotunda

RS

A
Dec 2003
Chamidae (jewel boxes)
Chama pacifica

Pseudochama corbieri

IP, RS

RS

E

A

Dec 2003

Jan 2005
Cardiidae (cockles)
Fulvia australis

Fulvia fragilis

Afrocardium richardi

IP, RS

WIO, RS

RS

E

E

E

Jan 2005

Jan 2005

Dec 2003
Tellinidae (tellins)
Tellina valtonis

Psammotreta praerupta

IO, RS

IP, RS

E

A

Dec 2003

Jan 2005
Psammobiidae (sanguin clams)
Hiatula ruppelliana

RS

E
Jan 2005
Mactridae (surf clams)
Mactra olorina

Mactra lilacea

RS

IP, RS

E

E

Dec 2003

Dec 2003
Mesodesmatidae
Atactodea glabrata

IP, RS

A
Dec 2003
Veneridae (venus clams)
Gafrarium pectinatum

Circenita callipyga

Clementia papyracea

Paphia textile

Mercenaria mercenaria

Ruditapes philippinarum

Antigona lamellaris

Dosinia erythraea

Timoclea roemeriana

IP, RS

IO, RS

IP, RS

IP, RS

AA

WP

IP, RS

RS, WP

RS

E

A

E

E

E

E

A

A

E

Jan 2005

Dec 2003

Jan 2005

Jan 2005

Dec 2003

Dec 2003

Jan 2005

Jan 2005

Jan 2005
Petricolidae
Petricola pholadiformis

AA

A
Dec 2003
Myidae (soft-shell clams)
Sphenia rueppelli

Mya arenaria

RS

AA

A

E

Dec 2003

Dec 2003
Gastrochaenidae (rock-borer clams)
Gastrochaena cymbium

IP, RS

E
Jan 2005
BIVALVIA - ANOMALODESMATA
Laternulidae (lantern clams)
Laternula anatina

IP, RS

E
Jan 2005
POLYPLACOPHORA
Chitonidae (chitons)
Chiton hululensis
RS, IP
A
Dec 2003
©ciesm 2002